1978.4-1982.3   University of Tokyo, Dept. of Science., Bachelor

1982.4-1984.3   Osaka University, Dept. of Medicine, Master course

1984,4-1988,4 Kyoto University, Dept. of Kyoto, Retained doctor thesis(1988,4)

1988.4-1990.9    Research scientist: Tokyo University, Dept. of Medicine

1990.10-1993.3 Research scientist: Basel(Swiss) University, BioCenter

1993.4-1995.7    Assistant professor: Kyoto Univ. Center for molecular genetics

1995.8-  1997,3 lecturer: Kyoto Univ. dept. of medicine

1997.4-2002.3     professor: Tokushima University, Dept of Arts and Science

2002,3-2003,10 Team Leader: Riken Center for Developmental Biology

2003,11- Professor: Nagoya Univerity, Dept. of Science

1997 Beckman Prize for encouragemente