1979-1980 Ochanomizu University(Japan)
　Undergraduate
　Population Genetics
   (Supervisor, S. Ishiwa-Chigusa)

1980-1982 Ochanomizu University(Japan)
　Graduate student
　Population Genetics
   (Supervisor, S. Ishiwa-Chigusa)

1982-1983 University of　Sussex(UK)
　Graduate student
　Population Biology
   (Supervisor, B. Charlesworth)

1983　National Cancer Center Research Institute (Japan)
　Graduate student
　Molecular Biology
   (Supervisor, K. Shimonohno)

1983-1988　Washington University(USA)
　Graduate student
　Genetics
   (Supervisor, R. H. Waterston)

1989-1998　Kyushu University (Japan)
　Assistant Professor
　Molecular Genetics

1998-2004　Nagoya University (Japan)
　Associate Professor
　Molecular Neurobiology

2004-         Nagoya University(Japan)
　Professor
　Molecular Neurobiology

1990  Nissan Science Foundation Fellowship, Japan

1990  Brain Science Foundation Fellowship, Japan

1996  The Genetics Society of Japan Young Investigator Award

2006　Saruhashi Award

2006　Inoue Prize for Science